For me, Always On as a Limitation has always implied that the Power causes some significant impediment to the normal functioning of the character. Force Field is a classic example of this for published Champions characters, since detrimental features are easy to define for it. Since a basic Force Field is always Visible while it's on, the character would always draw attention, have difficulty disguising himself, have trouble moving stealthily, etc. He might not be able to touch many objects or take in food. Those negative factors would be sufficient to warrant a Limitation IMO.
OTOH what you're describing for Mental Defense and Power Defense are rare and unusual circumstances. Most people don't use Telepathy to communicate for example, so an inability to lower Mental Defense would be an unusual circumstance (unless you're postulating an alien society where Telepathy is common, making the character the equivalent of a deaf person in hearing society). Similarly, people with Healing are uncommon, and the character can still heal normally.
Personally I don't think I would allow a Limitation on those special Defenses; however, depending on how often the inability to lower them would hinder the character, I might allow a small Physical Limitation, "Can't lower X Defense".
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